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Organism
  Animalia
      Arthropoda
            Insecta
              Pterygota
                    Coleoptera
                      Polyphaga
                          Scarabaeoidea
                            Scarabaeidae
                              Aphodiinae
                                     [Aphodiidae]
                                Aphodiini
                                     Aidophus
                                             Aidophus panamensis
                                                   [Aphodius panamensis]
                                             Aidophus parcus
                                                   [Aphodius knausii]
                                     Aphodius
                                             Aphodius ataeniodes
                                             Aphodius distinctus
                                             Aphodius fimetarius
                                             Aphodius hubbelli
                                             Aphodius lanei
                                             Aphodius lividus
                                             Aphodius luteolus
                                             Aphodius ruricola
                                             Aphodius tenuistriatus
                                             Aphodius tenvistriatus
                                             Aphodius vittatus
                                     Gonaphodioides
                                             Gonaphodioides chapini
                                                   [Aphodius chapini]
                                     Xenoheptaulacus
                                             Xenoheptaulacus tricostatus
                                                   [Xenoheptaulacus carinator]
                                Eupariini
                                     Ataenius
                                             Ataenius convexus
                                             Ataenius gracilis
                                             Ataenius imbricatus
                                             Ataenius panamensis
                                             Ataenius pearlensis
                                             Ataenius picinus
                                             Ataenius punctifrons
                                             Ataenius strigatus
                                             Ataenius wenzeli
                                     Auperia
                                             Auperia capitosa
                                     Batesiana
                                             Batesiana tuberculata
                                     Euparia
                                             Euparia castanea
                                     Euparixia
                                             Euparixia isthmia
                                     Euparixoides
                                             Euparixoides cribratus
                                     Lomanoxoides
                                             Lomanoxoides nigrolineatus
                                                   [Ataenius nigrolineatus]
                                Rhyparini
                                     Rhyparus
                                             Rhyparus blantoni
                                             Rhyparus isidroi
                                             Rhyparus sculpturatus
                                             Rhyparus spangleri
                                             Rhyparus zayasi
                              Aphodinae
                              Cetoniinae
                                     Balsameda
                                             Balsameda pulverulenta
                                Cetoniini
                                     Euphoria
                                             Euphoria basalis
                                             Euphoria candezei
                                             Euphoria children
                                             Euphoria eximia
                                             Euphoria fulgida
                                             Euphoria herbacea
                                             Euphoria histrionica
                                             Euphoria holochloris
                                             Euphoria inda
                                             Euphoria kerni
                                             Euphoria lesueuri
                                             Euphoria limbata
                                             Euphoria precaria
                                             Euphoria pulchella
                                             Euphoria sepulchralis
                                             Euphoria steinheili
                                             Euphoria testacea
                                             Euphoria yucateca
                                                   [Euphoria submetallica]
                                     Trichiotinus
                                             Trichiotinus assimilis
                                             Trichiotinus lunulatus
                                             Trichiotinus piger
                                             Trichiotinus texanus
                                     Genuchinus
                                             Genuchinus muzo
                                             Genuchinus nevermanni
                                     Giesbertiolus
                                             Giesbertiolus ornatus
                                Trichiini
                                     Dialithus
                                             Dialithus magnificus
                                             Dialithus scintillans
                                  Osmodermatina
                                     Inca
                                             Inca clathratus
                                               Inca clathratus sommeri
                                     Trigonopeltastes
                                             Trigonopeltastes delta
                                             Trigonopeltastes geometrica
                                             Trigonopeltastes sallaei
                                Valgini
                                     Valgus
                                             Valgus canaliculatus
                              Cetoninae
                                Gymnetini
                                     Allorhina
                                             Allorhina scabriuscula
                                     Amithao
                                             Amithao albopictum
                                             Amithao decemguttatus
                                             Amithao lafertei
                                                   [Amithao sparsus]
                                             Amithao spence
                                     Argyripa
                                             Argyripa anomala
                                             Argyripa subfasciata
                                             Argyripa subviolacea
                                     Cotinis
                                       Cotinis (Cotinis)
                                             Cotinis (Cotinis) lebasi
                                             Cotinis lebasi
                                             Cotinis mutabilis
                                             Cotinis nitida
                                     Guatemalica
                                             Guatemalica hueti
                                  Gymnetina
                                     Desicasta
                                             Desicasta reichei
                                     Marmarina
                                             Marmarina maculosa
                                     Gymnetis
                                             Gymnetis bajula
                                             Gymnetis chevrolati
                                             Gymnetis coturnix
                                             Gymnetis difficilis
                                             Gymnetis flaveola
                                             Gymnetis flavomarginata
                                             Gymnetis hebraica
                                             Gymnetis meleagris
                                             Gymnetis pantherina
                                             Gymnetis pardalis
                                             Gymnetis ramulosa
                                             Gymnetis stellata
                                             Gymnetis vandepolli
                                     Hadrosticta
                                             Hadrosticta viridiflua
                                     Hoplopyga
                                             Hoplopyga liturata
                                             Hoplopyga ocellata
                                     Chlorota
                                             Chlorota limbaticollis
                                     Cuemida
                                             Cuemida intermedia
                              Dinastinae
                              Dynastinae
                                Agaocephalini
                                     Aegopsis
                                             Aegopsis curvicornis
                                     Spodistes
                                             Spodistes armstrongi
                                             Spodistes batesi
                                             Spodistes hopei
                                             Spodistes mniszechi
                                Cyclocephalini
                                     Ancognatha
                                             Ancognatha gracilis
                                             Ancognatha humeralis
                                             Ancognatha manca
                                             Ancognatha scarabaeoides
                                             Ancognatha vexans
                                             Ancognatha vulgaris
                                     Aspidolea
                                             Aspidolea fuliginea
                                             Aspidolea kuntzeni
                                             Aspidolea notaticollis
                                             Aspidolea singularis
                                             Cyclocephala kuntzeni
                                     Cyclocephala
                                             Cyclocephala acuta
                                             Cyclocephala almitana
                                             Cyclocephala amazona
                                               Cyclocephala amazona subsp. amazona
                                                     [Cyclocephala beaumonti]
                                             Cyclocephala amazonica
                                             Cyclocephala amblyopsis
                                                   [Cyclocephala ambliopsis]
                                             Cyclocephala ampliata
                                             Cyclocephala atripes
                                             Cyclocephala brevis
                                             Cyclocephala brittoni
                                             Cyclocephala carbonaria
                                             Cyclocephala castaniella
                                                   [Cyclocephala obscurata]
                                             Cyclocephala complanata
                                             Cyclocephala concolor
                                             Cyclocephala confusa
                                             Cyclocephala conspicua
                                             Cyclocephala discicollis
                                             Cyclocephala discolor
                                             Cyclocephala epistomalis
                                             Cyclocephala erotylina
                                             Cyclocephala fasciolata
                                             Cyclocephala fulgurata
                                             Cyclocephala fulvipennis
                                             Cyclocephala gravis
                                             Cyclocephala guttata
                                             Cyclocephala herteli
                                                   [Cyclocephala barroensis]
                                             Cyclocephala hirta
                                             Cyclocephala isthmiensis
                                             Cyclocephala kaszabi
                                                   [Cyclocephala kazabi]
                                             Cyclocephala krombeini
                                             Cyclocephala labidion
                                             Cyclocephala ligyrina
                                             Cyclocephala lunulata
                                             Cyclocephala lurida
                                             Cyclocephala macrophylla
                                             Cyclocephala mafaffa
                                             Cyclocephala maffafa
                                             Cyclocephala melanae
                                             Cyclocephala melanocephala
                                                   [Dichromina ocularis]
                                             Cyclocephala mustacha
                                             Cyclocephala mutata
                                             Cyclocephala nigerrima
                                             Cyclocephala nigritarsis
                                             Cyclocephala nike
                                             Cyclocephala ovulum
                                                   [Cyclocephala ovalum]
                                             Cyclocephala pan
                                             Cyclocephala pardolocarnoi
                                             Cyclocephala perconfusa
                                             Cyclocephala porioni
                                             Cyclocephala prolongata
                                             Cyclocephala pubescens
                                             Cyclocephala quadripunctata
                                             Cyclocephala rogezi
                                             Cyclocephala sanguinicollis
                                             Cyclocephala santaritae
                                             Cyclocephala sexpunctata
                                                   [Stigmalia costaricana]
                                             Cyclocephala signaticollis
                                             Cyclocephala sinaloae
                                             Cyclocephala sinaloe
                                             Cyclocephala sororia
                                             Cyclocephala sparsa
                                                   [Cyclocephala landini]
                                             Cyclocephala spilopyga
                                             Cyclocephala stictica
                                             Cyclocephala stockwelli
                                             Cyclocephala suturalis
                                             Cyclocephala tridentata
                                             Cyclocephala variabilis
                                             Cyclocephala verticallis
                                             Cyclocephala weidneri
                                             Cyclocephala zodion
                                             Euetheola brevis
                                     Discinetus
                                     Dyscinetus
                                             Dyscinetus dubius
                                                   [Discinetus dubius]
                                             Dyscinetus laevipunctatus
                                             Dyscinetus morator
                                     Eriosceles
                                     Erioscelis
                                             Erioscelis columbica
                                                   [Eriosceles colombica]
                                             Erioscelis sobrina
                                     Mimeoma
                                             Mimeoma acuta
                                             Mimeoma englemani
                                     Mimeona
                                             Mimeona acuta
                                     Stenocrates
                                             Stenocrates bicarinatus
                                             Stenocrates carinatus
                                             Stenocrates cultor
                                             Stenocrates hardyi
                                             Stenocrates holomelanus
                                             Stenocrates laevicollis
                                             Stenocrates popei
                                Dynastini
                                     Dynastes
                                             Dynastes hercules
                                             Dynastes tityus
                                     Golofa
                                             Golofa costaricensis
                                             Golofa hirsuta
                                             Golofa imbelis
                                             Golofa imperialis
                                             Golofa obliquicornis
                                             Golofa solisi
                                     Magasoma
                                             Magasoma elephas
                                     Megasoma
                                             Megasoma actaeon
                                             Megasoma elephas
                                             Megasoma janus
                                Oryctini
                                     Coelosis
                                             Coelosis biloba
                                     Enema
                                             Enema endymion
                                             Enema pan
                                     Heterogomphus
                                             Heterogomphus chevrolati
                                             Heterogomphus dilaticollis
                                             Heterogomphus flohri
                                             Heterogomphus mniszechi
                                                   [Heterogomphus mnizechi]
                                             Heterogomphus philocteles
                                             Heterogomphus rugicollis
                                             Heterogomphus schoenherri
                                     Ligyrus
                                             Ligyrus bituberculatus
                                             Ligyrus ebenus
                                             Ligyrus gibbosus
                                             Ligyrus gyas
                                             Ligyrus morisnamer
                                             Ligyrus sallaei
                                     Megaceras
                                             Megaceras morpheus
                                             Megaceras septentrionis
                                     Podischnus
                                             Podischnus agenor
                                     Strategus
                                             Strategus aloeus
                                                   [Strategus gaillardi]
                                             Strategus antaeus
                                             Strategus hipposiderus
                                             Strategus jugurtha
                                             Strategus mormon
                                             Strategus oblongus
                                             Strategus simson
                                     Xyloryctes
                                             Xyloryctes furcata
                                             Xyloryctes jamaicensis
                                             Xyloryctes lobicollis
                                             Xyloryctes splendidus
                                             Xyloryctes telephus
                                Pentodontini
                                     Aphonus
                                             Aphonus brevicruris
                                             Aphonus texanus
                                             Aphonus tridentatus
                                     Barutus
                                             Barutus hartmanni
                                     Bothynus
                                             Bothynus complanus
                                             Bothynus medon
                                             Bothynus quadridens
                                     Euetheola
                                             Euetheola bidentata
                                             Euetheola humilis
                                     Eutheola
                                             Eutheola bidentata
                                             Eutheola humilis
                                     Orizabus
                                             Orizabus clunalis
                                     Oxygrylius
                                             Oxygrylius ruginasus
                                     Oxyligyrus
                                             Oxyligyrus zoilus
                                     Parapucaya
                                             Parapucaya amazonica
                                     Pucaya
                                             Pucaya castanea
                                     Tomarus
                                             Tomarus bituberculatus
                                             Tomarus ebenus
                                             Tomarus fossor
                                             Tomarus gyas
                                             Tomarus nasutus
                                             Tomarus similis
                                Phileurini
                                     Amblyodus
                                             Amblyodus taurus
                                     Amblyoproctus
                                             Amblyoproctus centroamericanus
                                     Archophileurus
                                             Archophileurus bullatus
                                             Archophileurus cribrosus
                                     Epiphileurus
                                             Epiphileurus cribatus
                                     Goniophileurus
                                             Goniophileurus femoratus
                                     Hemiphilerus
                                             Hemiphilerus laevicauda
                                     Hemiphileurus
                                             Hemiphileurus curoei
                                             Hemiphileurus cylindroides
                                                   [Hemiphileurus costaricensis]
                                             Hemiphileurus dyscritus
                                             Hemiphileurus nebulohylaeus
                                             Hemiphileurus panamanius
                                             Hemiphileurus pygidiopunctissimus
                                             Hemiphileurus simplex
                                             Hemiphileurus variolosus
                                     Homophileurus
                                             Homophileurus quadrituberculatus
                                             Homophileurus tricuspis
                                     Palaeophileurus
                                             Palaeophileurus panamensis
                                     Paraphileurus
                                             Paraphileurus venezuelensis
                                     Phileurus
                                             Phileurus carinatus
                                             Phileurus didymus
                                             Phileurus limicauda
                                             Phileurus truncatus
                                             Phileurus valgus
                                             Phileurus youngi
                                     Phyleurus
                                             Phyleurus didymus
                                     Hadrostictia
                                             Hadrostictia viridiflua
                                     Heterosternus
                                             Heterosternus oberthueri
                                             Heterosternus oberthuri
                              Hybosorinae
                                     Aporolaus
                                             Aporolaus fimbriatus
                                     Coilodes
                                             Coilodes castanea
                                Hybosorini
                                     Hybosorus
                                             Hybosorus illigeri
                                     Leucothyreus
                                             Leucothyreus femoratus
                              Melolonthinae
                                Diplotaxini
                                     Liogenys
                                             Liogenys macropelma
                                             Liogenys quadridens
                                     Diplotaxis
                                             Diplotaxis frondicola
                                             Diplotaxis muricata
                                             Diplotaxis poropyge
                                             Diplotaxis subcostata
                                             Diplotaxis zeteki
                                Hopliini
                                     Hoplia
                                             Hoplia festiva
                                             Hoplia squamifera
                                Macrodactylini
                                     Barybas
                                             Barybas microcrania
                                             Barybas oxygenys
                                     Ceraspis
                                             Ceraspis centralis
                                             Ceraspis nitida
                                             Ceraspis squamulifera
                                     Dichelonyx
                                             Dichelonyx albicollis
                                             Dichelonyx linearis
                                     Faula
                                             Faula centralis
                                     Isonychus
                                             Isonychus hirsutus
                                             Isonychus marmoratus
                                             Isonychus ocellatus
                                             Isonychus pictus
                                     Macrodactylus
                                             Macrodactylus solisi
                                             Macrodactylus suavis
                                     Plectris
                                             Plectris integrata
                                             Plectris pavida
                                     Pseudoserica
                                             Pseudoserica micans
                                                   [Plectris micans]
                                Melolonthini
                                     Hypothyce
                                             Hypothyce mixta
                                     Hypotrichia
                                             Hypotrichia spissipes
                                     Podolasia
                                             Podolasia bottimeri
                                Rhizotrogini
                                     Chlaenobia
                                             Chlaenobia panamana
                                     Phyllophaga
                                             Phyllophaga arcta
                                             Phyllophaga bowditchi
                                             Phyllophaga chiriquina
                                             Phyllophaga corrosa
                                             Phyllophaga crassisima
                                             Phyllophaga crenulata
                                             Phyllophaga cushmani
                                             Phyllophaga densata
                                             Phyllophaga drakii
                                             Phyllophaga ephilida
                                             Phyllophaga forsteri
                                             Phyllophaga foxii
                                             Phyllophaga fusca
                                             Phyllophaga futilis
                                             Phyllophaga gigantea
                                             Phyllophaga gracilis
                                             Phyllophaga hirsuta
                                             Phyllophaga hirticula
                                             Phyllophaga hirtiventris
                                             Phyllophaga ignava
                                             Phyllophaga implicita
                                             Phyllophaga inversa
                                             Phyllophaga invisa
                                             Phyllophaga lanceolata
                                             Phyllophaga lissopyge
                                             Phyllophaga micans
                                             Phyllophaga opaca
                                             Phyllophaga parvidens
                                             Phyllophaga profunda
                                             Phyllophaga prunina
                                             Phyllophaga pusillidens
                                             Phyllophaga quercus
                                             Phyllophaga renodis
                                             Phyllophaga rugosa
                                             Phyllophaga setifera
                                             Phyllophaga soror
                                             Phyllophaga submucida
                                             Phyllophaga vehemens
                                             Phyllophaga vicina
                                     Trichesthes
                                             Trichesthes calculisternis
                                                   [Phyllophaga calculisternis]
                                Sericini
                                     Astaena
                                     Serica
                                             Serica paralella
                                             Serica porcula
                                             Serica vespertina
                              Orphninae
                                Aegidiini
                                     Aegidium
                                             Aegidium asperatum
                                     Phalangochaeta
                                             Phalangochaeta tuberculata
                                     Psammodius
                              Rutelinae
                                Anomalini
                                     Anomala
                                             Anomala cincta
                                             Anomala clathrata
                                             Anomala colonica
                                             Anomala cupricolis
                                             Anomala cupricollis
                                             Anomala inconstans
                                             Anomala innuba
                                             Anomala marginata
                                             Anomala megalia
                                                   [Anomala megallia]
                                             Anomala nitidula
                                             Anomala parvula
                                             Anomala plurisulcata
                                             Anomala semitonsa
                                             Anomala specularis
                                             Anomala sticticoptera
                                             Anomala sulcans
                                             Anomala valdecostata
                                             Anomala valida
                                               Hydrangea anomala subsp. petiolaris
                                  Anomalina
                                     Moroniella
                                             Moroniella nitidula
                                     Paranomala
                                       Paranomala (Paranomala)
                                             Paranomala (Paranomala) discoidalis
                                                   [Anomala discoidalis]
                                             Paranomala (Paranomala) ochroptera
                                                   [Anomala ochroptera]
                                     Callistethus
                                             Callistethus fuscorubens
                                             Callistethus granulipygus
                                             Callistethus lativittis
                                             Callistethus marginatus
                                             Callistethus mimeloides
                                             Callistethus ruteloides
                                             Callistethus vanpatteni
                                Anomalinii
                                     Strigoderma
                                             Strigoderma arboricola
                                             Strigoderma marginata
                                             Strigoderma nodulosa
                                             Strigoderma orbicularis
                                             Strigoderma pygmaea
                                             Strigoderma vestita
                                Anoplognathini
                                     Phalangogonia
                                             Phalangogonia sperata
                                     Platycoelia
                                             Platycoelia grandicula
                                             Platycoelia humeralis
                                             Platycoelia mesosternalis
                                     Bolax
                                             Bolax magna
                                             Bolax magnus
                                             Bolax vittatus
                                     Chrysophora
                                             Chrysophora chrysochlora
                                     Epichalcoplethis
                                             Epichalcoplethis granulata
                                     Geniates
                                             Geniates flaviventris
                                             Geniates panamensis
                                             Geniates spinolae
                                Geniatini
                                     Trizogeniates
                                             Trizogeniates foveicollis
                                             Trizogeniates vittata
                                     Spodochamys
                                             Spodochamys cupreola
                                     Spodochlamys
                                             Spodochlamys cupreola
                                Rutelini
                                  Anticheirina
                                     Platyrutela
                                             Platyrutela cribrata
                                     Antichira
                                             Antichira isthmica
                                     Calomacraspis
                                             Calomacraspis haroldi
                                             Calomacraspis splendens
                                     Chasmodia
                                             Chasmodia amazona
                                             Chasmodia badia
                                             Chasmodia circumdata
                                     Chrysima
                                     Chrysina
                                             Chrysina argenteola
                                                   [Plusiotis argenteola]
                                             Chrysina aurigans
                                             Chrysina aurora
                                             Chrysina batesi
                                                   [Plusiotis batesi]
                                             Chrysina beraudi
                                             Chrysina boucardi
                                             Chrysina bruyeai
                                             Chrysina chalcothea
                                                   [Plusiotis chalcothea]
                                             Chrysina chrysargyrea
                                                   [Plusiotis chrysargyrea]
                                             Chrysina chrysopedila
                                                   [Plusiotis chrysopedila]
                                             Chrysina cunninghami
                                                   [Plusiotis cunninghami]
                                             Chrysina eyai
                                             Chrysina gloriosa
                                                   [Plusiotis gloriosa]
                                             Chrysina limbata
                                             Chrysina macropus
                                                   [Chrysina macropis]
                                             Chrysina magnistriata
                                                   [Plusiotis magnistriata]
                                             Chrysina marginata
                                                   [Plusiotis marginata]
                                             Chrysina optima
                                                   [Plusiotis optima]
                                             Chrysina oreicola
                                             Chrysina ratcliffei
                                                   [Plusiotis ratcliffei]
                                             Chrysina resplendens
                                                   [Plusiotis resplendens]
                                             Chrysina tricolor
                                             Chrysina woodi
                                                   [Plusiotis woodi]
                                     Cnemida
                                             Cnemida intermedia
                                  Heterosternina
                                     Macropoides
                                             Macropoides crassipes
                                     Lagochile
                                             Lagochile chiriquina
                                             Lagochile sparsa
                                     Macraspis
                                             Macraspis chrysis
                                             Macraspis costaricensis
                                             Macraspis hirtiventris
                                             Macraspis lucida
                                             Macraspis melanaria
                                             Macraspis tetradactyla
                                             Macraspis trifida
                                     Paracotalpa
                                             Paracotalpa deserta
                                             Paracotalpa puncticollis
                                     Parastasia
                                             Parastasia brevipes
                                     Pelidnota
                                             Pelidnota belti
                                             Pelidnota centroamericana
                                             Pelidnota chiriquina
                                             Pelidnota costaricensis
                                             Pelidnota frommeri
                                             Pelidnota glabra
                                             Pelidnota laevissima
                                             Pelidnota notata
                                             Pelidnota pallidipermis
                                             Pelidnota parallela
                                             Pelidnota polita
                                             Pelidnota prasina
                                             Pelidnota prolixa
                                             Pelidnota prommeri
                                             Pelidnota punctata
                                             Pelidnota punctulata
                                             Pelidnota strigosa
                                             Pelidnota virescens
                                             Pelidnota vitrescens
                                             Pelidnota xanthopya
                                             Pelidnota xanthopyga
                                     Plusiotis
                                     Popillia
                                             Popillia complanata
                                             Popillia japonica
                                     Pseudomacraspis
                                             Pseudomacraspis affinis
                                     Ptenomela
                                             Ptenomela gratiosa
                                     Rutela
                                             Rutela lineola
                                             Rutela pygidialis
                                             Rutela sanguinolenta
                                             Rutela viridiaurata
                              Scarabaeinae
                                Ateuchini
                                     Pedaridium
                                             Pedaridium bottimeri
                                             Pedaridium pilosum
                                     Bdelyropsis
                                             Bdelyropsis newtoni
                                     Bdelyrus
                                             Bdelyrus planovultum
                                             Bdelyrus seminudus
                                Canthonini
                                     Canthochilum
                                             Canthochilum taino
                                     Cryptocanthon
                                             Cryptocanthon denticulatum
                                             Cryptocanthon lobatus
                                     Melanocanthon
                                             Melanocanthon bispinatus
                                             Melanocanthon nigricornis
                                     Scybalocanthon
                                             Scybalocanthon moniliatus
                                Coprini
                                     Ateuchus
                                             Ateuchus aeneomicans
                                             Ateuchus agenor
                                             Ateuchus candezei
                                             Ateuchus guatemalensis
                                             Ateuchus histeroides
                                     Canthidium
                                             Canthidium angusticeps
                                             Canthidium ardens
                                             Canthidium aurifex
                                             Canthidium centrale
                                             Canthidium elegantulum
                                             Canthidium elegantum
                                             Canthidium haroldi
                                             Canthidium hespenheidei
                                             Canthidium leucopterum
                                             Canthidium pallidoa
                                             Canthidium pallidoalatum
                                                   [Canthidium pallidoaltum]
                                             Canthidium planovultum
                                             Canthidium tenebrosum
                                             Canthidium tuberifrons
                                             Canthidium variolosum
                                             Canthidium vespertinum
                                     Copris
                                             Copris boucardi
                                             Copris costaricensis
                                             Copris howdeni
                                             Copris incertus
                                             Copris lugubris
                                             Copris minutus
                                             Copris remotus
                                     Dichotomius
                                             Dichotomius agenor
                                             Dichotomius amicitiae
                                             Dichotomius annae
                                             Dichotomius carolinus
                                             Dichotomius femoratus
                                             Dichotomius gamboaensis
                                                   [Dichotomus gambocensis]
                                             Dichotomius satanas
                                     Dichotomus
                                     Ontherus
                                               Ontherus (C.) pseudodydimus
                                       Ontherus (Ontherus)
                                             Ontherus (Ontherus) azteca
                                                   [Ontherus strius]
                                             Ontherus brevipennis
                                             Ontherus coscineus
                                             Ontherus didymus
                                                   [Ontherus alexis]
                                             Ontherus pseudodidymus
                                     Oxysternom
                                     Oxysternon
                                             Oxysternon conspicillatum
                                                   [Oxysternom conspicillatum]
                                             Oxysternon silenus
                                             Oxysternon zikani
                                     Scatimus
                                             Scatimus ovatus
                                                   [Scatimus oratus]
                                             Scatimus plebejus
                                             Scatimus quadridentatus
                                     Uroxys
                                             Uroxys bidentatis
                                             Uroxys bidentis
                                             Uroxys boneti
                                                   [Uroxys macrocularis]
                                             Uroxys depressifrons
                                             Uroxys gatunensis
                                             Uroxys gorgon
                                             Uroxys metagorgon
                                             Uroxys microcularis
                                             Uroxys micros
                                             Uroxys platypyga
                                Eurysternini
                                     Eurysternus
                                             Eurysternus caribaeus
                                             Eurysternus claudicans
                                             Eurysternus foedus
                                             Eurysternus magnus
                                             Eurysternus mexicanus
                                             Eurysternus plebejus
                                             Eurysternus velutinus
                                Oniticellini
                                     Euoniticellus
                                             Euoniticellus intermedius
                                Onitini
                                     Phanaeus
                                             Phanaeus adonis
                                             Phanaeus beltianus
                                                   [Phanaeus belfianus]
                                             Phanaeus chalcomelas
                                             Phanaeus difformis
                                             Phanaeus endymion
                                             Phanaeus hermes
                                             Phanaeus howdeni
                                             Phanaeus meleagris
                                             Phanaeus pyrois
                                             Phanaeus tridens
                                             Phanaeus vindex
                                     Onthocharis
                                             Onthocharis panamensis
                                Onthophagini
                                     Onthophagus
                                             Onthophagus acuminatus
                                             Onthophagus atriglabrus
                                             Onthophagus atriglabus
                                             Onthophagus atrosericeus
                                             Onthophagus batesi
                                             Onthophagus belorhinus
                                             Onthophagus chryses
                                             Onthophagus coscineus
                                             Onthophagus crinitus
                                             Onthophagus cyclographus
                                             Onthophagus dicranius
                                             Onthophagus gazella
                                             Onthophagus gazellinus
                                             Onthophagus incensus
                                             Onthophagus lebasi
                                             Onthophagus limonensis
                                             Onthophagus marginicollis
                                             Onthophagus medorensis
                                             Onthophagus mirabilis
                                             Onthophagus nuchicornis
                                             Onthophagus nyctopus
                                             Onthophagus orphnoides
                                             Onthophagus panamensis
                                             Onthophagus praecellens
                                             Onthophagus sharpi
                                             Onthophagus stockwelli
                                             Onthophagus transisthmius
                                Phanaeini
                                     Coprophanaeus
                                             Coprophanaeus chiriquensis
                                             Coprophanaeus ohausi
                                             Coprophanaeus telamon
                                     Sulcophanaeus
                                             Sulcophanaeus carnifex
                                             Sulcophanaeus cupricollis
                                             Sulcophanaeus noctis
                                             Sulcophanaeus noctius
                                             Sulcophanaeus velutianus
                                Scarabaeini
                                     Canthon
                                             Canthon aequinoctia
                                             Canthon aequinoctiale
                                             Canthon aequinoctialis
                                             Canthon angustatus
                                             Canthon cyanellus
                                             Canthon hartmanni
                                             Canthon humboldti
                                             Canthon humbolti
                                             Canthon imitator
                                             Canthon indigaceus
                                             Canthon juvencus
                                             Canthon lamprimus
                                             Canthon laprimus
                                             Canthon literatus
                                             Canthon maculatum
                                             Canthon meridionalis
                                             Canthon moniliatus
                                             Canthon morsei
                                             Canthon mutabilis
                                             Canthon sallei
                                             Canthon septemmaculatum
                                             Canthon septemmaculatus
                                             Canthon subhyalinus
                                             Canthon vazquezae
                                             Canthon vigilans
                                     Deltochilum
                                             Deltochilum acropyge
                                             Deltochilum gibbosum
                                             Deltochilum howdeni
                                             Deltochilum mexicanum
                                             Deltochilum pseudoparile
                                             Deltochilum scabriusculum
                                             Deltochilum seminudus
                                             Deltochilum valgum
                                Sisyphini
                                     Sisyphus
                                             Sisyphus mexicanus
                                     Stephanorrhina
                                             Stephanorrhina guttata
                                     Sulcophaneus
                                             Sulcophanaeus velutinus
                                     Thyriochlorota
                                             Thyriochlorota chibchana